Behavioural experiments are usually carried out to study the abilities of all animals. To this end, the animals are trained using the method of operant conditioning. The key to this training method lies in the positive reinforcement of desired behaviours. When the animal displays the desired behaviour, the correct response is first signalled by a short whistle from the training whistle (secondary reinforcer). The whistle is then followed by food (primary reinforcer). This forms the basis for the behavioural experiments, which are usually carried out as detection or discrimination experiments.